New material of Sinopsocus oligovenus Lin, 1976 (Hemiptera, Sternorrhyncha, Protopsyllidiidae) with preserved body structures from the Middle Jurassic Daohugou Bed, northeastern China

Abstract

Protopsyllidioidea is an extinct superfamily of Sternorrhyncha (Hemiptera) known from the late Carboniferous to the late Cretaceous (Hakim et al., 2019; Boderau et al. 2026). It can be distinguished from other hemipterans by the combination of a rather narrow area between C and ScP + R, the absence or scarcity of crossveins, especially in the distal half of the forewing, a simple RP, a long common stem M + CuA emerging from R + M + CuA, the presence of strong setae on all main veins, and the fusion of CuA with M to form a common stem M + CuA. This combination of characters provides a more reliable basis for distinguishing the taxon from other hemipterans (Becker-Migdisova, 1985; Klimaszewski, 1995; Grimaldi, 2003; Nel et al., 2012; Huang et al., 2022). Currently, five families are recognized within Protopsyllidioidea: Permopsyllidiidae Becker-Migdisova, 1985, Protopsyllidiidae Carpenter, 1931, Postopsyllidiidae Hakim et al., 2019, Paraprotopsyllidiidae Hakim et al., 2021 and Lovedopsyllididae Boderau et al., 2024.